The narrative, over-seen by Hirohiko Araki, takes place immediately following the events of Stardust Crusaders. Jotaro Kujo and his allies, who had just defeated the evil vampire DIO, are suddenly attacked by friends and enemies who have either disappeared or died along the way. It is then they are approached by a young Robert E. O. Speedwagon, who possesses a piece of the Saint's Corpse that allows him to travel through time and space as well as vanquish the evil influence possessing the others. He leads Jotaro and his allies on a journey through time, space, and alternate universes, in order to stop another iteration of DIO and his disciple Father Enrico Pucci from obtaining the other pieces of the Saint's Corpse in DIO's quest to finally be free of the Joestar family and achieve his own personal heaven.[3] Fighters from all 8 parts of JoJo's Bizarre Adventure appear as Jotaro travels through time to meet his great-great grandfather Jonathan Joestar, his grandfather Joseph Joestar from a point in his youth, Joseph’s illegitimate son Josuke Higashikata, DIO's son (biologically Jonathan’s) Giorno Giovanna, and his own daughter Jolyne Cujoh, while Jonathan and Josuke meet their alternate universe counterparts, the paraplegic jockey Johnny Joestar and the amnesiac only known as "Josuke Higashikata."

Once the heroes finish their tasks, they gather the Corpse Parts they have, but notice one is missing. Suddenly, the Joestars are all transported before a DIO with a powered-up version of The World known as The World: Over Heaven. The World: Over Heaven easily repels the Joestars' attacks, even managing to overcome the reality-nullifying power of Giorno's Gold Experience Requiem and the infinite rotation of Johnny's Tusk ACT 4 before using his Corpse's part to draw all the other Corpse parts to him. Before DIO can eliminate the Joestars, however, they manage to escape, as DIO realizes Jotaro managed to keep a Corpse's part to himself. After they were transported, Funny Valentine appears to the group and reveals that DIO couldn't take Jotaro's Corpse Part because their Stands are still the same type, even after the former's evolution. Elaborating further, he explains that when Johnny had captured him with Tusk ACT 4's infinite rotation, he used D4C to escape to a parallel Earth of the original universe in which DIO won the events of Part 3, evolved his Stand, and conquered the world, becoming a godlike being known as Heaven Ascension DIO. After Valentine's unsuccessful attempt to bring Heaven Ascension DIO over to his reality to destroy Johnny, DIO decided to invade the main world so he could conquer it. After Valentine reveals DIO's plan to the group, they split up in order to reclaim the Saint Corpse Parts from DIO's underlings once more, retrieve the remaining allies still being manipulated, and find out the secret of DIO's evolved Stand. Meanwhile, Valentine repeats his vow to protect his nation before entering a hopeless battle with DIO, in which he is defeated and erased from existence.

After accomplishing the first two goals but unable to figure out the secret behind DIO's Stand, the JoJos and Speedwagon return to Egypt to confront DIO, who is defeated, but absorbs most of the Joestars and their allies' souls, as well as the Corpse parts. DIO rewrites the universe to his own liking, but Jotaro and Jolyne survive to face him. DIO quickly overpowers them both and takes Jolyne's soul in the process, leaving Jotaro on his own. DIO manages to land a punch with The World Over Heaven and attempts to erase Jotaro only for it to fail. Jotaro reveals that Star Platinum, being the same type of Stand as The World Over Heaven, has attained Over Heaven as well just as it had attained The World's time stopping abilities before and stopped his erasure with it. With the playing field levelled Jotaro remembers Valentine's warning of two iterations of the same person destroying each other upon contact. Jotaro reveals that he has obtained prime DIO's armlets, which fuses with Heaven Ascension DIO and badly damages his hands, the source of The World Over Heaven's ability to rewrite reality. Without his reality-warping powers, Jotaro gains the upper hand and destroys Heaven Ascension DIO in a manner much like in the main universe. After DIO's defeat, all realities are restored and Jotaro returns to meeting his allies at the end of Part 3. However, the credits reveal that prior to giving up Star Platinum: Over Heaven, Jotaro rewrote his timeline so that everyone who died in Parts 1-6 survived, including all of his own friends who had died, and it's also revealed that he did the same for the Parts 7-8 timeline.


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