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In the main story, Android 16 plays a prominent role - having mysteriously been revived, along with a "mysterious army" of Super Androids who look like the Z Fighters, and then the Z Fighters are taken down left and right. Android 21, an all-new female character, also plays a major role in the story.

There are three story scenarios: the Super Warrior Arc where Goku is the main character, the Enemy Warrior Arc where Frieza is the main character and the Android 21 Arc, where Androids 18 and 21 are the main characters. Each arc features a similar but somewhat different story. The main plot of the game is revealed in the Android 21 Arc.

The stories of this game occurs sometime after the Universe 6 Saga story-line but before the "Future" Trunks Saga. This is shown by Bulma and Android 18 appearing as they did in Dragon Ball Z: Resurrection ‘F’ and its anime adaption Golden Frieza Saga, while Goku and Vegeta are wearing their regular gi/armor. Gohan also notes that Frieza had returned before the events of the game and Vegeta mentions that Gohan has recently begun training under Piccolo again but has not yet shown the results of his training. Sorbet, although he never appears in the story, is depicted with five fingers in the graphic that shows when his Bad Ring Laser attack is triggered when Frieza fights. This is something the minor antagonist displayed only through the anime adaptation, while his official design only had four fingers in the original movie, as well as its limited manga release. Frieza mentions Piccolo's sacrifice to save Gohan from him, Gotenks when asked by Frieza about Future Trunks, claimed that he doesn't know him and Piccolo is amazed that he has to team up with Frieza, something he thought would never happen. When talking with Cell, Goku notes that he has never been to Hell before. In the Super Warriors Arc, when rescuing Good Buu, Goku notes that Buu is just as asleep as he was during the written test required to participate in the Tournament of Destroyers. The other characters in the beginning of the Super Warriors Arc react as if this is the first time that Goku has his body possessed by someone else.

The gameplay borrows concepts from several other fighting games, primarily the Marvel vs. Capcom series' control scheme and team mechanics, with three primary attack buttons and one unique action button plus a few others. Unlike the Arc System Works-developed Extreme Butōden, FighterZ plays out in a scheme much closer to many other of the traditional Arc System Works-developed fighting games (via running on the same graphics as Guilty Gear Xrd). The three primary attack buttons unlike in other fighting games come with a twist; light attacks are the same as in other games, while medium attacks are a mix of both traditional medium and heavy attacks, while a heavy attack is a move that knocks the opponent away with a heavy blow. The unique action button by itself, or with a direction or special move motion allows for a ki blast or ki-based special moves to be performed.

Players each select three characters to form a team. One character is controlled, and can be switched with one of the other characters at any time. Players can also call one of their other characters to perform an "Assist" move, allowing simultaneous attacks and combos with the entire team. All three of a team's characters must be defeated for a match to end.

The universal "Vanish" and "Sparking Blast" moves resemble the Roman Cancel system commonly found in Arc System Works games and the Guilty Gear series' Dust attacks respectively (styled akin to both before Xrd and after Xrd), while the "Super Dash" move resembles the Homing system of Arcana Heart and is akin to the Homing dashes seen in other Dragon Ball fighting games. Just like in the said classic Dragon Ball fighting games, characters can "Ki Charge" to raise their power level akin to other fighting game mechanics that allow for gauge-charging. Also added are the ability to deflect Ki blasts or other projectiles via a parry mechanic, as well as being able to use a Dragon Rush that acts akin to a basic throw that launches the victim away while the attack flight-dashes after them.

Other mechanics involve the usage of the auto-combo system utilized by fighting games as of Persona 4: Arena (tap one button for an automatic preset combo), which now also applies to the medium attack button as well (giving all characters two auto-combos). Forward + medium attack is also a universal overhead attack for all characters, which cannot be chained out of.

Nearly all characters are also planned to have differing playstyles/archetypes apart from each other akin to more contemporary fighting games as opposed to using recycled base movesets such as in the Budokai series.

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