In the C 63 S, the four-cylinder unit develops 350 kW (476 hp) at 6725 rpm. This makes it the world's most powerful series-production four-cylinder engine. The maximum torque of 545 Nm is available at 5250‑5500 rpm. A belt-driven starter generator (RSG) combines an alternator and starter in one component. The RSG starts the combustion engine and ensures the basic supply of the ancillary components such as air conditioning or driving lights. This also applies when the vehicle is at a traffic light and the charge level of the high-voltage battery is no longer sufficient to support the low-voltage on-board electrical system. The RSG is integrated into the 400-volt high-voltage network.

Power is transmitted via the AMG SPEEDSHIFT MCT 9G transmission (MCT = Multi-Clutch Transmission), in which a wet start-off clutch replaces the torque converter. It reduces weight and, thanks to its lower inertia, optimises response to accelerator pedal input, especially during spurts and load changes. The elaborately calibrated software ensures extremely short shift times as well as fast multiple downshifts if required. In addition, the intermediate throttle function in the "Sport" and "Sport+" driving modes delivers a particularly emotionally appealing gearshift experience. There is also the RACE START function, which ensures optimal acceleration from a standstill.

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