When the player reaches the Shadowlord's Castle, it is revealed that both Devola and Popola are androids working for Project Gestalt. The twins are revealed to have lived for thousands of years as a result of the initial experiments of the project, and have thus been charged with keeping eye on other experiments as a result: namely over the members of the entire party, including Yonah.

They were made to provide support when the day has come for Gestalts to return to the Replicants' vessels. They are tasked with extracting data from Gestalts' souls, making new Replicants' bodies, and returning a dead Replicant's data to its Gestalt.

NieR Replicant Playlist
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