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Earth is governed by the United Citizen Federation, a military government founded generations earlier by "veterans" after democracy and social scientists brought civilization to the brink of ruin. Citizenship is gained only through federal service, which grants rights—such as to vote and to breed—that are withheld from ordinary civilians. Humans, who are now spacefaring, conduct colonization missions throughout the galaxy, bringing them into conflict with a race of highly evolved insectoid creatures dubbed "Arachnids" or, derisively, "bugs".

Despite the protestations of his parents about the risks, teenage jock Johnny Rico enlists as a mobile infantryman to remain close to his girlfriend, spaceship pilot Carmen Ibanez. Their psychic friend Carl Jenkins joins Military Intelligence, while Isabelle "Dizzy" Flores, who is in love with Rico, deliberately transfers to his squad. Carmen eventually breaks up with Rico because their career paths separate them and she has developed a mutual attraction with fellow pilot Zander Barcalow. During training, Rico impresses drill sergeant Zim, earning a promotion to squad leader. He makes a mistake during a training exercise causing the death of one of his squad and the resignation of another, resulting in his demotion and flogging.

An invasion force is deployed to Klendathu, the Arachnids' home planet, but the military intelligence underestimates the Arachnids' defensive abilities, leading to thousands of casualties. Badly wounded, Rico is rescued by lieutenant Jean Rasczak, his former high-school teacher, but is mistakenly reported dead, devastating Carmen. Following his recovery, Rico, Dizzy, and squadmate Ace Levy join Rasczak's special forces unit, the "Roughnecks". Rico eventually earns the rank of Corporal for his feats against the Arachnids, including killing a gigantic "Tanker Bug" on Tango Urilla, and becomes romantically involved with Dizzy. The Roughnecks respond to a distress call on Planet P, where they reconnoitre an Arachnid-ravaged outpost but are quickly overwhelmed by the bugs. Carmen and Zander recover the surviving Roughnecks by dropship, but not before Dizzy is killed and Rico is forced to mercy-kill the mutilated Rasczak. The group returns to the fleet assembled in orbit above P, where Dizzy is eulogized.

Jenkins, now a high-ranking colonel, explains the Roughnecks were deliberately sent into the trap, justifying it as a necessary sacrifice to prove the existence of a "Brain Bug"; an intelligent Arachnid strategically directing the others. He grants Rico command of the Roughnecks and field-promotes him to lieutenant, ordering him back to P to capture the Brain Bug. As the battle commences, Carmen's ship is destroyed by the Arachnids, forcing her and Zander to flee in an escape shuttle that crash lands into an underground tunnel system. The pair are captured by the Arachnids and the Brain Bug uses its proboscis to consume Zander's brain, absorbing his knowledge. Rico sends his squad to complete their mission while he, Ace, and their squadmate Watkins find Carmen and hold the Arachnids at bay with a miniature nuclear bomb.

The Brain Bug escapes while the Arachnids attack and mortally wound Watkins, who sacrifices himself by detonating the bomb while his teammates escape. On the surface, they learn that Zim has captured the Brain Bug, and the assembled troops rejoice as Jenkins psychically detects it is afraid. A propaganda advertisement details how the Brain Bug is being invasively studied, supposedly to learn its secrets and ensure humanity's victory. The advert encourages the viewers to enlist and do their part in the war so they can become like Carmen, now captain of her own ship, and Rico, who enthusiastically leads his troops into another battle.

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