In this part we discover that Cortana threatened Lord Hood and the UNSC, in turn threatened Earth. Cortana wanted Earth to surrender. If Earth governments wants to fight, she'll make sure they'd lose. Cortana also went to Atriox, leader of the Banished and asked the same thing. He of course refused, so she blew up his home planet of Doisac.

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In 2559, the UNSC Infinity is attacked and boarded by the Banished. Atriox (Ike Amadi) quickly defeats the Master Chief (Steve Downes) and hurls him into space. Six months later, a UNSC pilot code-named Echo 216 (Nicolas Roye) recovers the Chief adrift in space. While Echo-216 wants to flee, Master Chief insists on continuing the fight against the Banished. After destroying a Banished dreadnaught in space, the Chief and Echo-216 travel to Zeta Halo, which has been severely damaged by an unknown calamity. On the ring, Master Chief locates the Weapon (Jen Taylor), an artificial intelligence designed to imitate Cortana (Jen Taylor) to capture her for deletion. While the Weapon states that she was successful in her mission, she has failed to self-delete per her programming, for unknown reasons. Master Chief retrieves the Weapon, and begins to experience visions of Cortana's memories and thoughts left as residual data following her death.

Zeta Halo is controlled by the Banished, now led by Atriox's lieutenant Escharum (Darin De Paul) after Atriox's apparent death. Master Chief, the Weapon, and a reluctant Echo-216 rally scattered UNSC forces on the ring's surface. After infiltrating a Banished installation known as "The Tower", Master Chief learns from a dying Spartan that the Banished have excavated a Forerunner facility known as the Conservatory. Inside, Master Chief encounters Despondent Pyre, the AI caretaker of Zeta Halo. Despondent Pyre attempts to warn Master Chief of a threat contained on the installation, but is subsequently destroyed by the Harbinger, an alien being reawakened by the Banished upon their arrival on Zeta Halo. The Harbinger claims that her people, the Endless, survived the firing of the Halo Array, but were subsequently imprisoned by the Forerunners on Zeta Halo. She has allied herself with the Banished in order to rebuild the Silent Auditorium, a facility which will allow her to free the Endless.

Echo-216 reveals that he was an engineer aboard the Infinity who stole a Pelican to flee the battle. The Chief comforts him, admitting his own guilt over his failure to save Cortana. Master Chief and the Weapon stop the Halo's repair process by disabling a series of spires on the ring. When the Harbinger attempts to hack the Weapon, the Chief activates a failsafe to delete the AI. The Weapon repels the Harbinger and deactivates the failsafe, angrily asking why Master Chief does not trust her. After seeing a vision in which Cortana uses the Guardians to destroy the Brutes' home planet of Doisac, the Weapon realizes that she is a copy of Cortana, and insists that Master Chief delete her so she does not become like her predecessor; Master Chief refuses, stating that he wants to trust her. Echo-216 is captured by Escharum, who is revealed to be terminally ill, and brought to the House of Reckoning, the headquarters of the Banished, as bait for Master Chief. Unwilling to let Echo-216 die, Master Chief infiltrates the facility and kills Escharum.

Master Chief and the Weapon infiltrate the Silent Auditorium to stop the Harbinger. The two learn that Cortana was captured by Atriox after the Weapon locked her down; upon realizing that Atriox intended to use the ring, she destroyed herself and damaged Zeta Halo while also preventing the Weapon from being deleted so that she could continue to assist the Master Chief in Cortana's place. A final message left behind by a remorseful Cortana says goodbye to the Chief, urging him and the Weapon not to repeat her mistakes and to become stronger from them. The Chief defeats the Harbinger, but he is unable to stop her from contacting someone regarding the Endless. Afterwards, the two reunite with Echo-216, resolving to continue to fight against the Banished presence on the ring. Echo-216 reveals that his real name is Fernando Esparza, and Master Chief tells the Weapon to choose a name for herself, as Cortana had done before her. She says that she has decided on the perfect name for herself, but what it is isn't revealed.

In a post-credits scene, Atriox uses a key to unlock a sealed door, leading to the Endless. If the game is finished on Legendary difficulty, the scene opens with the date 97,368 BCE in which Despondent Pyre is heard speaking with the Forerunner Grand Edict about the imprisonment of the Endless. When Despondent Pyre states that she cannot oversee the study of the Endless alone, the Grand Edict replies that the AI Offensive Bias has been deployed to assist her.

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